参考文献 References
[1] Azadeh H. 2013. Stylar polymorphism, reciprocity and incompatibility systems in Nymphoides montana (Menyanthaceae) endemic to southeastern Australia. Plant Systematics and Evolution 299: 389-401.
[2] Sánchez J M, Ferrero V, Arroyo J, et al. 2010. Patterns of style polymorphism in five species of the South African genus Nivenia (Iridaceae). Annals of Botany, 106: 321–331.
[3] Rodrigues EB, Consolaro H. 2013. Atypical distyly in Psychotria goyazensis Mull. Arg. (Rubiaceae), an intramorph self-compatible species. Acta Botanica Brasilica, 27: 155–161.
[4] Watanabe K, Shimizu A, Sugawara T. 2014. Dioecy derived from distyly and pollination in Psychotria rubra (Rubiaceae) occurring in the Ryukyu Islands, Japan. Plant Species Biology, 29:181–191.
[5] Barrett SCH, Jesson LK, Baker AM. 2000. The evolution and function of stylar polymorphisms in flowering plants. American Journal of Botany, 85: 253–265.
[6] 周伟, 王红. 2009. 被子植物异型花柱及其进化意义. 植物学报, 44: 742–751.
[7] Faivre A E, McDade L A. 2001. Population-level variation in the expression of heterostyly in three species of Rubiaceae: Does reciprocal placement of anthers and stigmas characterize heterstyly?. American Journal of Botany, 88: 841–853.
[8] Kissling J, Barrett SCH .2013.Variation and evolution of herkogamy in Exochaenium(Gentianaceae): Implications for the evolution of distyly. Annals of Botany, 112: 95–102.
[9] Aronne G, Buonanno M, Micco VD. 2014. Assessment of distyly syndrome in Primula palinuri Petagn. a rare species living on maritime vertical cliffs. Plant Systematics and Evolution, 300: 917–924.
[10] Liu WL, Wang YF, Chen Q, et al. 2014. Pollination of invasive Eichhornia crassipes (Pontederiaceae) by the introduced honeybee (Apis mellifera L.) in South China. Plant Systematics and Evolution. 299:817–825.
[11] Wang Y, Wang QF, Guo YH, et al. 2005. Reproductive consequences of interactions between clonal growth and sexual reproduction in Nymphoides peltata: a distylous aquatic plant. New Phytologist, 165: 329–336.
[12] Ma YP, Xie WJ, Tian XL, Sun WB, Wu ZK, Milne R. 2014. Unidirectionalhy bridization and reproductive barriers between two heterostylous primrose species in north-west Yunnan, China. Annals of Botany, 113: 763–775.
[13] Barrett SCH. 2002. Mating strategies in flowering plants: the outcrossing-selfing paradigm and beyond. The Royal Society of London Series B Biological Sciences, 358: 991–1004.
[14] Barrett SCH. 2019. ‘A most complex marriage arrangement’: recent advances on heterostyly and unresolved questions. New Phytologist224: 1051–1067.
[15] Charlesworth D, Charlesworth B. 1979. A model for the evolution of distyly. American Naturalist, 114: 467–498.
[16] Webb CJ, Lloyd DG .1986. The avoidance of interference between the presentation of pollen and stigmas in angiosperms. II. Herkogamy. New Zeeland Journal of Botany. 24:163–178.
[17] Richards AJ 1998. Lethal linkage and its role in the evolution of plant breeding systems. In: Owens S.J., Rudall PJ.(eds), Reporductive Bioloyg in Systematies, Consevration, and Eeonomic Botnay, 71-83. Royal Botanic Garden, Kew
[18] Pailler T, Humeau L, and Thomeson JD.1998. Distyly and heteromorphic incompatibility in oceanic island species of Erythroxylum (Erythroxylaceae). Plant Systematics and Evolution, 213: 187–198.
[19] Zhong L, Barrett SC, Wang XJ, Wu ZK, Sun HY, Li DZ et al. (2019) Phylogenomic analysis reveals multiple evolutionary origins of selfing from outcrossing in a lineage of heterostylous plants. N Phytol 224:1290–1303
[20] Richards, AJ. 2002. Primula. London: BT Batsford.
[21] Barrett SCH, Shore JS (2008) New Insights on Heterostyly: Comparative Biology,Ecology and Genetics. In: Franklin-Tong VE (ed) Self-Incompatibility in Flower-ing Plants: Evolution, Diversity, and Mechanisms. Springer, Berlin, p 3–32
[22] Mora-Carrera E, Stubbs R, Keller B, Léveillé-Bourret T, Conti E (2023) Different mole-cular changes underlie the same phenotypic transition: origins and con-sequences of independent shifts to homostyly within species. Mol Ecol 32:61–78
[23] Huu CN, Keller B, Conti E, Kappel C, Lenhard M (2020) Supergene evolution via stepwise duplications and neofunctionalization of a floral-organ identity gene.Proc Natl Acad Sci USA 117:23148–23157
[24] Barrett SCH. 2019. ‘A most complex marriage arrangement’: recent advances on heterostyly and unresolved questions. New Phytologist224: 1051–1067.
[25] Zhang W, Hu YF, He X, Zhou W, Shao JW (2021) Evolution of Autonomous Selfing in Marginal Habitats: Spatiotemporal Variation in the Floral Traits of the Distylous Primula wannanensis. Front Plant Sci 12:781281
[26] Almeida NM, Castro CC, Leite AV, Novo RR, Machado IC. 2013. Floral polymorphism in Chamaecrista flexuosa (Fabaceae-Caesalpinioideae): A possible case of atypical enantiostyly? Annals of Botany, 112:1117–1123.
[27] Hodgins, KA, Barrett SCH. 2008. Natural selection on floral traits through male and female function in wild populations of the heterostylous daffodil Narcissus triandrus. 62: Evolution,1751-1763.
[28] Brys R, Jacquemyn H (2015) Disruption of the distylous syndrome in Primula veris. AnnBot 115:27–39
[29] Richards AJ .1986. Plant breeding systems. George Allen & Unwin, London.
[30] Ishihama F, Washitani I. 2011. Post-pollination process in a partially self-compatible disylous plant, Primula sieboldii (Primulaceae). Plant Species Biology, 26: 213–220.
[31] Minuto L, Guerrina M, Roccotiello E, Roccatagliata N, Mariotti MG, Casazza G. 2014. Pollination ecology in the narrow endemic winter-flowering Primula allionii (Primulaceae). Journal of Plant Research, 127: 141–150.
[32] Ganders E. 1979. The biology of heterostyly. New Zealand Journal of Botany, 17: 607–635.
[33] Beach JH, Bawa KS. 1980. Role of pollinators in the evolution of dioecy from distyly. Evolution 34: 1138-1142.
[34] Cesaro AC, Barrett SCH, Maurice S, Vaissiere BE, Thompson JD. 2004. An experimental evaluation of self-interference in Narcissus assoanus: functional and evolutionary implications. Journal of Evolutionary Biology, 17: 1367–1376.
[35] Zhu XF, Jiang XF, Li L, Zhang ZQ, Li QJ. 2015. Asymmetrical disassortative pollination in a distylous primrose: the complementary roles of bumblebee nectar robbers and syrphid flies. Scientific Report. doi:10.1038/srep07721.
[36] Barrett SCH. 1992. Evolution and function of heterostyly. Berlin: Springer-Verlag
[37] Kulbaba, MW; Worley AC. 2008. Floral design in Polemonium brandegei (Polemoniaceae): Genetic and phenotypic variation under hawkmoth and hummingbird pollination. International Journal of Plant Sciences. 169:509-522.
[38] 张大勇. 2004. 植物生活史进化与繁殖生态学. 北京: 科学出版社: 96–180.
[39] Kitamoto N, Ueno S, TakenakaA, et al. 2006. Effect of flowering phenology on pollen flow distance and the consequences for spatial genetic structure within a population of Primula sieboldii (Primulaceae). American Journal of Botany, 93: 226–233.
[40] Wu ZK, Zhang CQ. 2010. Comparative study of pollination biology of two closely related alpine Primula species, namely Primula beesiana and P. bulleyana (Primulaceae). Journal of Systematics and Evolution, 48:109–117.
[41] Vanhoenacker D, Ågren J, Ehrlén J. 2006. Spatio-temporal variation in pollen limitation and reproductive success of two scape morphs in Primula farinosa. New Phytologist, 169: 615–621.
[42] Geert AV, Rossum FV, Triest L. 2010. Do linear landscape elements in farmland act as biological corridors for pollen dispersal? Journal of Ecology, 98, 178–187.
[43] Brys R, Jacquemyn H, Hermy M. 2007. Impact of mate availability, population size, and spatial aggregation of morphs on sexual reproduction in a distylous, aquatic plant. American Joural of Botany, 94: 119–127.
[44] Kitamoto N, Ueno S, Tsumura Y, et al. 2008. Effect of population density of compatible neighbours on inbreeding level within a Primula sieboldii population. Ecological Research, 23: 307–315.
[45] Guggisberg A, Mansion G, Kelso S, et al. 2006. Evolution of biogeographic patterns, ploidy levels, and breeding systems in a diploid-polyploid species complex of Primula. New Phytologist 171: 617-632.